An on-leave army officer named Kate (Julia Roberts) meets Holden (Bradley Cooper), a gay man, on a flight from Iraq to Los Angeles. The owner of a florist shop, Reed (Ashton Kutcher) proposes to his girlfriend, Morley (Jessica Alba), while learning his best friend, Julia (Jennifer Garner), has a boyfriend named Harrison (Patrick Dempsey) who turns out to be married. An office assistant named Liz (Anne Hathaway) is dating the mailroom assistant, Jason (Topher Grace). Julia's mother, Estelle (Shirley MacLaine) is a happy retiree who must reveal an affair from long ago to her husband, Edgar (Hector Elizondo). Tyler Harrington (Taylor Lautner) and Samantha Kenny (Taylor Swift) are high schoolers who are deeply in love. Meanwhile, a publicist named Kara (Jessica Biel) is seemingly alone with no date on Valentine's Day working for her boss Kelvin Briggs (Jamie Foxx) whom she falls in love with working together. All these casually intertwining stories of the heart take place over the course of one Valentine's Day.
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