Giselle (Amy Adams) lives in Andalasia, an animated fairy tale world with talking animals, characters breaking into song and "happily ever after" a foregone conclusion. Prince Edward (James Marsden), her designated true love saves her from a troll and they plan to get married the next day. Queen Narissa (Susan Sarandon), Edward's evil stepmother, schemes to protect her throne. She throws Giselle into a portal that transports her to a world without "happily ever afters" -- modern (and live-action) New York City.
In New York, Giselle soon meets Robert Philip (Patrick Dempsey), a hardened divorce lawyer. At his daughter (Rachel Covey) Morgan's insistence, he lets Giselle stay at his apartment. Giselle, following the ways of her fairy tale world, recruits urban animals—pigeons, cockroaches and rats to clean his apartment while she makes a gown out of his curtains. Robert's fiancĂ©, Nancy (Idina Menzel), misunderstands the situation and begins to argue with Robert.
He decides to part with Giselle at Central Park, but rejoins her after seeing her give the money he gave her to an old woman. During their walk through Central Park, Giselle questions Robert on how he displays his affection for Nancy and spontaneously starts the musical production number "That's How You Know" with many performers in the park joining her. Giselle helps Robert reconcile with Nancy by sending an apology on Robert's behalf, along with tickets to the King's and Queen's Ball.
Meanwhile, Queen Narissa's henchman Nathaniel (Timothy Spall) follows Edward and Pip, a speaking chipmunk who is friends with Giselle, who have journeyed to New York to save Giselle. They stop at a motel, where Nathaniel questions his relationship with Narissa after watching a soap opera. He sneaks out to give Giselle a poisoned apple. Pip is unable to speak coherently in this world and has a frustrating time alerting the Prince of the minion's intentions. When Nathaniel fails twice to poison Giselle, Narissa becomes infuriated.
As Giselle and Robert spend more time together, Giselle discovers that the real world is much more complicated than she realized, while Robert is affected by her optimism and idealism. Edward continues to look for Giselle and eventually finds her at Robert's apartment. While Edward is eager to take Giselle home, they go on a date around New York at her insistence. To Nancy's chagrin, Giselle and Edward attend the King's and Queen's Ball. After Nancy and Edward pair off to dance, Giselle dances with Robert. During their dance, Giselle realizes that Robert is her true love as he, very softly, sings the lyrics of the song to her. Unbeknownst to them, Narissa has traveled to New York from Andalasia. Under her old hag's disguise, she manages to successfully poison Giselle, but is stopped by Edward before she can escape with Giselle's unconscious body.
A remorseful Nathaniel reveals Narissa's plot, admits his deeds and reveals that the spell of the poison apple has to be broken before midnight or Giselle will die. Robert realizes that only true love's kiss can revive Giselle, but it doesn't work for Edward. Desperate, Edward begs a hesitant Robert to try. Nancy gives her consent and Robert kisses the sleeping Giselle. Giselle awakens to Robert's kiss. Narissa uses the distracting moment to break free. She transforms into a dragon, deciding to write her own ending to the story. Taking Robert hostage, she lures Giselle out the window and up to the top of the Woolworth Building. With help from Pip, Narissa falls from the roof to her death, exploding into magic dust at street level. Giselle catches Robert, and they manage to keep themselves from falling off the roof. Edward and Robert are praised as heroes.
Nancy goes with Edward to Andalasia and marries him. Giselle opens a boutique in New York City, where she is assisted by both humans and animals. Both Nathaniel (in New York) and Pip (in Andalasia) become successful authors. The last scene shows Giselle, Robert, and Morgan playing together and living happily ever after as a family.
In New York, Giselle soon meets Robert Philip (Patrick Dempsey), a hardened divorce lawyer. At his daughter (Rachel Covey) Morgan's insistence, he lets Giselle stay at his apartment. Giselle, following the ways of her fairy tale world, recruits urban animals—pigeons, cockroaches and rats to clean his apartment while she makes a gown out of his curtains. Robert's fiancĂ©, Nancy (Idina Menzel), misunderstands the situation and begins to argue with Robert.
He decides to part with Giselle at Central Park, but rejoins her after seeing her give the money he gave her to an old woman. During their walk through Central Park, Giselle questions Robert on how he displays his affection for Nancy and spontaneously starts the musical production number "That's How You Know" with many performers in the park joining her. Giselle helps Robert reconcile with Nancy by sending an apology on Robert's behalf, along with tickets to the King's and Queen's Ball.
Meanwhile, Queen Narissa's henchman Nathaniel (Timothy Spall) follows Edward and Pip, a speaking chipmunk who is friends with Giselle, who have journeyed to New York to save Giselle. They stop at a motel, where Nathaniel questions his relationship with Narissa after watching a soap opera. He sneaks out to give Giselle a poisoned apple. Pip is unable to speak coherently in this world and has a frustrating time alerting the Prince of the minion's intentions. When Nathaniel fails twice to poison Giselle, Narissa becomes infuriated.
As Giselle and Robert spend more time together, Giselle discovers that the real world is much more complicated than she realized, while Robert is affected by her optimism and idealism. Edward continues to look for Giselle and eventually finds her at Robert's apartment. While Edward is eager to take Giselle home, they go on a date around New York at her insistence. To Nancy's chagrin, Giselle and Edward attend the King's and Queen's Ball. After Nancy and Edward pair off to dance, Giselle dances with Robert. During their dance, Giselle realizes that Robert is her true love as he, very softly, sings the lyrics of the song to her. Unbeknownst to them, Narissa has traveled to New York from Andalasia. Under her old hag's disguise, she manages to successfully poison Giselle, but is stopped by Edward before she can escape with Giselle's unconscious body.
A remorseful Nathaniel reveals Narissa's plot, admits his deeds and reveals that the spell of the poison apple has to be broken before midnight or Giselle will die. Robert realizes that only true love's kiss can revive Giselle, but it doesn't work for Edward. Desperate, Edward begs a hesitant Robert to try. Nancy gives her consent and Robert kisses the sleeping Giselle. Giselle awakens to Robert's kiss. Narissa uses the distracting moment to break free. She transforms into a dragon, deciding to write her own ending to the story. Taking Robert hostage, she lures Giselle out the window and up to the top of the Woolworth Building. With help from Pip, Narissa falls from the roof to her death, exploding into magic dust at street level. Giselle catches Robert, and they manage to keep themselves from falling off the roof. Edward and Robert are praised as heroes.
Nancy goes with Edward to Andalasia and marries him. Giselle opens a boutique in New York City, where she is assisted by both humans and animals. Both Nathaniel (in New York) and Pip (in Andalasia) become successful authors. The last scene shows Giselle, Robert, and Morgan playing together and living happily ever after as a family.
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